This timeline of ceratosaur research is a chronological listing of events in the history of paleontology focused on the ceratosaurs, a group of relatively primitive, often horned, predatory theropod dinosaurs that became the apex predators of the southern hemisphere during the Late Cretaceous. The nature and taxonomic composition of the Ceratosauria has been controversial since the group was first distinguished in the late 19th century. In 1884 Othniel Charles Marsh described the new genus and species Ceratosaurus nasicornis from the Late Jurassic Morrison Formation of the western United States. He felt that it belonged in a new family that he called the Ceratosauridae. He created the new taxon Ceratosauria to include both the Ceratosauridae and the ostrich-like ornithomimids. The idea of the Ceratosauria was soon contested, however. Later that same decade both Lydekker and Marsh's hated rival Edward Drinker Cope argued that the taxon was invalid.

The idea of the Ceratosauria would regain some support more than thirty years later when Gilmore argued in its favor in 1920. Nevertheless, the validity of Ceratosauria was disputed throughout much of the 20th century by researchers like Romer, Lapparent, Lavocat, Colbert, and Charig. However, in 1986, more than a century after Marsh first coined the name, Jacques Gauthier revived the idea. Three years later, Rowe published a new definition of Ceratosauria, all taxa more closely related to Ceratosaurus than to birds, based on Gauthier's use of the term. This modern use of the term was thought to include the many theropods discovered since the 1880s known as coelophysoids. Ceratosaurus itself had loose joints between bones in the skull whose interpretation has been controversial. Paleontologist Robert T. Bakker has interpreted this condition as an adaptation to swallow prey larger than it would otherwise be able to fit through its jaws.

Since the 1980s, major developments in ceratosaur taxonomy have centered on the discovery of the Abelisauridae, a new family of large ceratosaurs that were among the dominant predators of the southern hemisphere during the Cretaceous. One of the most notable of these was Carnotaurus, an unusual horned theropod with a short face. More recent noteworthy non-abelisaur ceratosaur discoveries include the protruding-toothed noasaurid Masiakasaurus knopfleri, named after the lead guitarist from Dire Straits.

19th century

1880s

1884 Othniel Charles Marsh described the new genus and species Ceratosaurus nasicornis. He also erected the new family Ceratosauridae and named the Ceratosauria to include both Ceratosaurus and the Ornithomimidae.

1888 Richard Lydekker rejected the validity of the Ceratosauria and Ceratosauridae.

1890s

1892 Cope rejected the validity of the Ceratosauria and Ceratosauridae.

1896 Charles Depéret described the new species Megalosaurus crenatissimus, based mostly on vertebrae. Marsh described the new genus and species Labrosaurus sulcatus.

20th century

1900s

1901

Arthur Smith Woodward described the new genus and species Genyodectes serus.

1910s

1919 Othenio Abel rejected the validity of the Ceratosauria and Ceratosauridae.

1920s

1920 Werner Janensch described the new genus and species Elaphrosaurus bambergi. Charles Whitney Gilmore recognized the Ceratosauria and/or Ceratosauridae as valid.

1921

Charles William Andrews described the new genus and species Sarcosaurus woodi. Charles Alfred Matley described the new genus and species Lametasaurus indicus.

1925 Janensch described the new species Ceratosaurus roechlingi. He also described the new species Labrosaurus stechowi.

1930s

1931 Hem Chandra Das Gupta described the new genus and species Orthogoniosaurus matleyi.

1932 Von Huene described the new genus and species Laevisuchus indicus. He also described the new genus and species Coeluroides largus. He also described the new species Dryptosauruoides grandis. He also described the new genus and species Velocipes guerichi. He also described the new species Sarcosaurus andrewsi. Huene recognized the Ceratosauria and/or Ceratosauridae as valid.

1933 Von Huene and Matley described the new genus and species Indosaurus matleyi. They also described the new genus and species Indosuchus raptorius. They also described the new genus and species Jubbulpuria tenuis. They also described the new genus and species Ornithomimoides barasimlensis. They also described the new species O. mobilis. Alfred Sherwood Romer rejected the validity of the Ceratosauria and Ceratosauridae.

1950s

1955

René Lavocat erected the new genus Majungasaurus to house the species "Megalosaurus" crenatissimus, and designated a neotype specimen. Albert-Félix de Lapparent and Lavocat rejected the validity of the Ceratosauria and Ceratosauridae.

1956 Romer rejected the validity of the Ceratosauria and Ceratosauridae. Von Huene recognized the Ceratosauria and/or Ceratosauridae as valid.

1960s

1964 Colbert rejected the validity of the Ceratosauria and Ceratosauridae.

1966 Romer rejected the validity of the Ceratosauria and Ceratosauridae.

1970s

1970 Rodney Steel rejected the validity of the Ceratosauria and Ceratosauridae.

1979

Hans-Dieter Sues and Philippe Taquet described the new genus and species Majungatholus atopus. Alan Jack Charig rejected the validity of the Ceratosauria and Ceratosauridae.

1980s

1980 José Bonaparte and Jaime Powell described the new genus and species Noasaurus leali. They classified it in a new family, the Noasauridae.

1985

Bonaparte and Fernando Emilio Novas described the new genus and species Abelisaurus comahuensis. Bonaparte described the new genus and species Carnotaurus sastrei.

1986 Robert T. Bakker thought the loose joints in the skull of Ceratosaurus was an adaptation allowing it to swallow prey that would otherwise be too large. Ricardo Martínez and others described the new genus and species Xenotarsosaurus bonapartei. Gauthier "resurrected the name Ceratosauria" to apply to the sister group of the tetanuran theropods. However, while he listed the members of his Ceratosauria, he did not propose a formal phylogenetic definition.

1989 Rowe formally defined the Ceratosauria as theropods more closely related to Ceratosaurus nasicornis than birds, based on Gauthier's 1986 usage of the term.

1990s

1990 Rowe and Gauthier published a node-based definition of Ceratosauria.

1991 Jean Le Loeuff and Eric Buffetaut described the new genus and species Tarascosaurus salluvicus. Bonaparte described the new genus and species Velocisaurus unicus. Novas regarded the Ceratosauria as consisting of two sister groups, the Coelophysoidea and their sister group, the Neoceratosauria.

1994 Thomas Holtz named the Coelophysoidea, then thought to be ceratosaurs. He defined them as all theropods more closely related to Coelophysis than to Ceratosaurus. Holtz defined the Neoceratosauria as all theropods closer to Ceratosaurus than to Coelophysis. He also defined the ceratosaur subclade Abelisauroidea as all theropods closer to Carnotaurus sastrei than to Ceratosaurus nasicornis.

1995 Hugues Accarie and others described the new genus and species Genusaurus sisteronis.

1996 Bonaparte described the new genus and species Ligabueino andesi. Sankar Chatterjee and Dhiraj Kumar Rudra observed that abelisaur teeth had lower crowns than other carnivorous dinosaurs either within or outside of Ceratosauria.

1998

Coria and Leonardo Salgado described the new genus and species Ilokelesia aguagrandensis. Sereno redefined Ceratosauria as all neotheropods closer to Coelophysis bauri than to birds. However, this definition never received broad acceptance by the scientific community because the Rowe had already defined the group in 1989, and therefore had priority. He also split the Abelisauridae into two subfamilies; the Abelisaurinae (all abelisaurids more closely related to Abelisaurus than to Carnotaurus) and Carnotaurinae (all abelisaurids more closely related to Carnotaurus than to Abelisaurus).

Sampson and others described a more complete specimen of Majungatholus atops. They performed an X-ray CT scan of the skull of Majungatholus atopus. They found that the dome on the animal's skull formed of its frontal bone was actually hollow. This suggests that this structure was purely for display rather than used in fights between Majungatholus.

21st century

2000s

2000 Madsen and Welles described the new species Ceratosaurus dentisulcatus and C. magnicornis.

2001 Sampson, Carrano, and Forster described the new genus and species Masiakasaurus knopfleri. They noted that at least six individuals were preserved together. This makes Masiakasaurus the first neoceratosaur to be preserved in association with others of its species.

2002

Coria, Chiappe, and Dingus described the new genus and species Aucasaurus garridoi. Kellner and Campos described the new genus and species Pycnonemosaurus nevesi.

2003 Wilson and others described the new genus and species Rajasaurus narmadensis.

2004 Calvo, Rubilar-Rogers and Moreno described the new genus and species Ekrixinatosaurus novasi. P. C. Sereno, J. A. Wilson, and J. L. Conrad described the new genus and species Rugops primus as well as the new genus Spinostropheus.

2006 Malkani described the new genus and species Vitakridrinda sulaimani.

2007 Allain and others described the new genus and species Berberosaurus liassicus.

2008 Sereno and Brusatte described the new genus and species Kryptops palaios. Canale and others described the new genus and species Skorpiovenator bustingorryi.

2009 Xu and others described the new genus and species Limusaurus inextricabilis.

2010s

2010 M. D. Ezcurra, F. L. Agnolin, and Novas described the new genus and species Austrocheirus isasii. Novas and others described the new genus and species Rahiolisaurus gujaratensis.

2012 Pol and Rauhut described the new genus and species Eoabelisaurus mefi.

2013

Farke and Sertich described the new genus and species Dahalokely tokana. Tortosa and others described the new genus and species Arcovenator escotae.

2014 Sánchez-Hernández and Benton described the new genus and species Camarillasaurus cirugedae. Sebastian G. Dalman described the new genus and species Fosterovenator.

2016 Filippi and others described the new genus and species Viavenator exxoni.

2017 Longrich and others described the new genus and species Chenanisaurus barbaricus.

2018 Dal Sasso and others described the new genus and species Saltriovenator zanellai. Delcourt and Iori described the new genus and species Thanos simonattoi.

2019 Partially preserved ilium of an indeterminate abelisaur theropod is reported from the Upper Cretaceous Kem Kem Beds (Morocco) by Zitouni et al. (2019). Langer and others described the new genus and species Vespersaurus paranaensis.

2020s

2020 Cerronia and others described the new genus and species Tralkasaurus cuyi.

See also History of paleontology Timeline of paleontology Timeline of coelophysoid research

Footnotes

References

External links