Thermospermine synthase (, TSPMS, ACL5 (ACAULIS5), SAC51) is an enzyme characterised from Thalassiosira pseudonana and Arabidopsis thaliana with systematic name S-adenosylmethioninamine:spermidine 3-aminopropyltransferase (thermospermine synthesizing). It catalyses a chemical reaction in which an aminopropyl group is transferred from the coenzyme, S-adenosylmethioninamine to spermidine, giving thermospermine:
This enzyme is required for correct xylem specification through regulation of the lifetime of the xylem elements.
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