Gloydius is a genus of pit vipers endemic to Asia, also known as Asian moccasins or Asian ground pit vipers. Named after American herpetologist Howard K. Gloyd,Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). The Eponym Dictionary of Reptiles. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (Gloydius, p. 102). this genus is very similar to the North American genus Agkistrodon. 24 species are currently recognized.

Geographic range Species of Gloydius are found in Russia, east of the Ural Mountains through Siberia, Iran, Himalayas from Pakistan, India, Nepal, China, Korea, Japan and the Ryukyu Islands.

Species SpeciesTaxon authorSubsp.*Common nameGumprecht A, Tillack F, Orlov NL, Captain A, Ryabov S (2004). Asian Pitvipers. First Edition. Berlin: Geitje Books. 368 pp. .Geographic rangeG. angusticepsJ. Shi, Yang, Huang, Orlov & P. Li, 20180Zoige pit viperChina, (Sichuan, Qinghai, and Gansu).G. blomhoffii(H. Boie, 1826)0Japanese mamushiJapan.G. brevicauda(Stejneger, 1907)0Short-tailed mamushiNorth Korea, South Korea, and China.G. caraganus(Eichwald, 1831)0Karaganda pitviperUzbekistan, Turkmenistan, Tajikistan, Kyrgyzstan, and Kazakhstan.G. caucasicus(Nikolsky, 1916)0Caucasian pitviperTurkmenistan, Azerbaijan, Iran, and Afghanistan.G. chambensisKuttalam, Santra, Owens, Selvan, Mukherjee, Graham, Togridou, Bharti, Shi, Shanker & Malhotra, 2022 (Gloydius chambensis, new species).0 Chamba pitviperChamba District, Himachal Pradesh.G. changdaoensisL. Li, 19990China.G. cognatus(Gloyd, 1977)0Alashan pitviperChina.G. halysT(Pallas, 1776)4Siberian pitviperRussia, east of the Ural Mountains through Siberia, Iran, Mongolia to northern and central China, as well as the southern Ryukyu Islands.G. himalayanus(Günther, 1864)0Himalayan pitviperAlong the southern slopes of the Himalayas from northeastern Pakistan, to northern India (Kashmir, Punjab) and Nepal. Found at 1524–3048 m altitude.G. huangiK. Wang, Ren, Dong, Jiang, Shi, Siler & Che, 20190Lancang Plateau viperTibet.G. intermedius(Strauch, 1868)0Central Asian pitviperSoutheastern Azerbaijan, northern Iran, southern Turkmenistan, northwestern Afghanistan, southern Russia, northwestern China and Mongolia.G. lateralisZhang, S. Shi, C. Li, Yan, P. Wang, Ding, Du, Plenković-Moraj, Jiang & J. Shi, 20220Jiuzhaigou County, China. (Gloydius lateralis, new species).G. lipipengiJ. Shi, J-C. Liu & Malhotra, 20210Nujiang pit viperChina.G. liupanensisN. Liu, Z. Song & Luo, 19890Gansu, China.G. monticola(F. Werner, 1922)0Likiang pitviperThe mountains of northern Yunnan in China.G. qinlingensis(M. Song & F. Chen, 1985)0Shaanxi, China.G. rickmersiWagner, Tiutenko, Borkin & Simonov, 20150KyrgyzstanG. rubromaculatusJ. Shi, P.Li & J. Liu, 20170Tibetan Plateau.G. saxatilis(Emelianov, 1937)0Rock mamushiGloyd HK, Conant R (1990). Snakes of the Agkistrodon Complex: A Monographic Review. Society for the Study of Amphibians and Reptiles. 614 pp., 52 plates. LCCN 89-50342. .Russia (eastern Siberia), northeastern China and North and South Korea.G. shedaoensis(Zhao, 1979)0Shedao island pitviperShedao Island, off the coast of Liaotung, China.G. stejnegeri(Rendahl, 1933)0Gobi pitviperChina and Mongolia.G. strauchi(Bedriaga, 1912)0Strauch's pitviperThe Tibetan Plateau in the provinces of Tsinghai and western Szechwan, China.G. swildJ. Shi & Malhotra, 20210China.G. tsushimaensis(Isogawa, Moriya & Mitsui, 1994)0Tsushima Island pitviperTsushima Island, Japan.G. ussuriensis(Emelianov, 1929)0Ussuri mamushiFar east Russia (Primorskiy Kray), northeastern China, North and South Korea, as well as Quelpart Island. (*) Not including the nominate subspecies. (T) Type species.

In the above list, a taxon author in parentheses indicates that the species was originally described in a different genus.

Taxonomy Due to the strong morphological similarity, these snakes were classified in the genus Agkistrodon until very recently. However, by 1999 cladistic studies clearly showed that Agkistrodon did not form a clade (indeed, it was not even paraphyletic) and was thus split into several genera.

A new species, G. tsushimaensis, was described by Isogawa, Moriya & Mitsui (1994). It is referred to as the Tsushima island pitviper and is found only on Tsushima Island, Japan.

References

Further reading

Hoge AR, Romano-Hoge SA (1981). "Poisonous Snakes of the World. I. Checklist of the Pitvipers: Viperoidea, Viperidae, Crotalinae". Memórias do Instituto Butantan 42/43: 179-309. (Gloydius, new genus). Isogawa, Kiyoshi; Moriya, Akira; Mitsui, Sadaaki (1994). "A new snake from the genus Agkistrodon (Serpentes: Viperidae) from Tsushima Island, Nagasaki Prefecture". Japanese Journal of Herpetology 15: 101-111. (Agkistrodon tsushimaensis, new species).

External links

Taxonomy comparison by Dr. Wolfgang Wüster at Dr. Brian Grieg Fry's Int'l Venom & Toxin Database. Accessed 25 July 2007.