The Carcharodini are a tribe in the skipper butterfly subfamily Pyrginae. They are a very diverse but quite plesiomorphic and inconspicuous group distributed throughout the tropics.Brower (2009)
Formerly, when only four tribes of Pyrginae were recognized, they were included in the Pyrgini, which at that time contained a massive number of genera. But the Pyrginae have since been reorganized to make them and their tribes monophyletic, leading most modern authors to treat the Carcharodini as distinct tribe. However, the old circumscription of the Pyrgini was by and large just as correct from a phylogenetic perspective.
The tribe of the Pyrgini sensu lato most closely related to the Pyrgini sensu stricto are the strikingly different Achlyodidini. As many consider it desirable to treat this lineage as a distinct tribe, the Carcharodini naturally need to be considered distinct too.
Genera These genera belong to the tribe Carcharodini:
Agyllia Grishin, 2020 Arteurotia Butler & H. Druce, 1872 Austinus Mielke & Casagrande, 2016 Bolla Mabille, 1903 Burca Bell & Comstock, 1948 Carcharodus Hübner, Clytius Grishin, 2019 Conognathus Felder & Felder, 1862 Cyclosemia Mabille, 1878 (eyed skippers) Ernsta Grishin, 2020 Favria Tutt, 1906 Fuscocimex Austin, 2008 Gomalia Moore, 1879 (marbled or African mallow skippers) Gorgopas Godman & Salvin, 1894 Hesperopsis Dyar, 1905 (sootywings) Iliana Bell, 1937 Incisus Grishin, 2019 Jera Lindsey, 1925 Mictris Evans, 1955 Mimia Evans, 1953 Morvina Evans, 1953 Muschampia Tutt, 1906 Myrinia Evans, 1953 Nisoniades Hübner, 1819 Noctuana E. Bell, 1937 Ocella Evans, 1953 Pachyneuria Mabille, 1888 Pellicia Herrich-Schaffer, 1870 Perus Grishin, 2019 Pholisora Scudder, 1872 (sootywings) Polyctor Evans, 1953 Sophista Plötz, 1879 Spialia Swinhoe, 1912 (grizzled skippers & sandmen) Staphylus Godman & Salvin, 1896 (scallopwings) Viola Evans, 1953 (Viola skippers) Viuria Grishin, 2019 Windia H. Freeman, 1969 Xispia Lindsey, 1925
Footnotes
References
(2009): Tree of Life Web Project – Carcharodini. Version of 2009-JUN-14. Retrieved 2009-DEC-26.